Following an education in Gonville and Caius College, Cambridge, William Sponne became Rector of Heavingham and Blofield in Norfolk, resigning the latter on being collated to the Archdeaconry of Norfolk on 21st December 1419. In June 1422 he became Rector of Towcester, a post which he held until the appointment of his successor, Thomas Taylard, in February 1447. Archdeacon Sponne died later in 1448.

His monument in the church is very similar to that of Bishop Richard Fleming of Lincoln who died 1431 in the same diocese. The monument is medieval and of a form in fashion at the time for higher ecclesiastics. It is a chest tomb, having a fully clothed effigy of the person on the top wearing a cassock, surplice and tippet, with fur-lined sleeves and collar. Beneath his effigy and inside the arches in the lower section is a representation of his emaciated corpse, indicative of mortality and evidently meant to teach humility. Cadaver tombs or table tombs were the fashion between 1420 and 1480, and surviving examples are rare, there being only about 150 still in existence. Whilst this design is rather unusual, it is the type of monument normally associated with large establishments, such as cathedrals and abbeys and to find one in a typical parish church such as St Lawrence's, clearly suggests that William Sponne was highly regarded in his own lifetime.
The monument is of clunch stone The sides of the tomb are open arches and on the ledge, the arms of Sponne are repeated. The head and hands are carved locally of oak with the face as a portrait; they were restored to the tomb in 1992 after being removed in 1884, lost and finally re-discovered in the attics of the Vicarage in the 1980s. At that time it was believed that the wooden appendages were themselves replacements, however radiocarbon dating in 1984 confirmed that the present wooden head is the original. The tinted prints inscribed '"Drawn on the spot" by T. Trotter' dated 1801 show clearly the wooden head attached to the tomb at that date. Over time the figure has been repainted several times and not always in the same colours. An article in the Sunday Telegraph tells of a disaster in the early 1980's when someone, not suitably qualified, repainted Archdeacon Sponne using "Humbrol" paint. Considerable restoration work was required to repair the damage.
The present position of the monument is not the original and a closely adjacent floor slab notes the following: "In a vault beneath this stone lie the remains of the Rev William Sponne Archdeacon of Norfolk whose monument was removed from this spot in November 1835 to make room for the new pewing of the Church. He died A D 1448". When this monument was removed from its original position in the chancel it was discovered that the skeleton was reportedly in a perfect state of preservation, lying on a bed of fine white sand. Within a few hours of the grave being opened the bones had crumbled to dust. The tomb has since been returned to its former position. This was supervised by Mr G Forsyth Lawson Church Architect in 1938. In 1440 Archdeacon Sponne bought a dwelling, known then as the Tabard, which changed its name in 1635 to The Talbot Inn, with adjoining lands. This was amongst his properties when he died and continued in the Charity's ownership until it was sold in 1895. It is now known at the Sponne Shopping Arcade.

The shield within the East window of Sponne's chapel is glass bought from The Tabard Inn, once owned by the charity. Shield of Sponne surrounded by fragments of green drapery and canopy work and the inscription Willi(a)m Sponn, in Gothic textura.
The drawing of the tomb near the top of this page was made by T.Trotter in 1801. His inscription at bottom reads:
"North View of the MONUMENT and CHANTRY CHAPEL of WILLm. SPONN, at TOWCESTER, Northamptonshire. At the upper end of the South ile was formerly a small Chapel parted off and called Sponns Chantry, of which this Monument now forms the North side, the above gives a view of the Chapel as it now appears, the Monument itself is still in a very high state of preservation, and may be look'd upon as a truly Valuable remnant of Antiquity, for which in all probability we are indebted to the very high respect his memory is held in, by every Inhabitant of the Town of Towcester. What the emaciated Skeleton like Figure underneath is intended to represent, the Artist cannot to a certainty assertain; in his peregrinations and enquiries he has discovered no less than Six of these horrible Figures, one at Winchester; two at Salisbury, one at Frome, one at Bristol, and the present at Towcester. They are uniformly attach'd to the Monuments of Ecclesiasticks, and an interesting Tradition usually repeated by those which shew them to straingers, to substantiate their personality, but it may be with more propriety be presumed that they are given as moral Lessons; as Emblems to remind us that frail humanity tho clad in the most gorgeous Apparel, tho incensed with adulation, and pampered with delicacies, to this Complexion it must come at last."
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